Non-human species may exhibit various stages in the evolution of morality.
Conceptualizing morality as a form of behavior opens the possibililty of observing it in other species. Indeed, if complex features evolve gradually, one might well expect to find stages of protomorality, incipient morality or various precursors in organisms besides humans. An important resource in understanding the evolution of humans and their culture, then, is comparative behavior. Even if the behavior is not strictly genetic, one may still find informative phylogenetic patterns or similarities based on common ancestry. Studies of primate behavior are potentially valuable. Transitions and intermediate stages may be more concretely envisioned or documented.